Publications

In Press

Brown, B.R.P., L.M. Khasoha, P. Lokeny, R.P. Jakopak, C.G. Reed, M. Dyck, A. Wambua, S.D. Newsome, T.M. Palmer, R.M. Pringle, J.R. Goheen, and T.R. Kartzinel. Spatiotemporal variation in the gut microbiomes of co-occurring wild rodent species. Ecosphere

Esmaeili, S., M-R Hemami, P. Kaczensky, K.A. Schoenecker, S.R.B. King, B. Shahriari, C. Walzer, and J.R. Goheen. Rainfall reduces the potential for competitive suppression of a globally endangered ungulate by livestock. Biological Conservation

Rezvani, A., M-R Hemami, J.R. Goheen, P. Kaczensky, S. Pourmanafi, S. Fakheran, and S. Esmaeili. Rethinking connectivity modeling for high-mobility ungulates: insights from a globally endangered equid. Landscape Ecology

2024

Kamaru, D.N., T.M. Palmer, C. Riginos, A.T. Ford, J. Belnap, R.M. Chira, J.M. Githaiga, B.C. Gituku, B.R. Hays, C.M. Kavwele, A.K. Kibungei, C.T. Lamb, N.J. Maiyo, P.D. Milligan, S. Mutisya, C.C. Ng'weno, M. Ogutu, A.G. Pietrek, B.T. Wildt, and J.R. Goheen. Disruption of an ant-plant mutualism shapes interactions between lions and their primary prey. Science 383:433-438. PDF

2023

Alston, J.M., D.A. Keinath, C.K.R. Willis, C.L. Lausen, J.M. O'Keefe, J.D. Tyburec, H.G. Broders, P.R. Moosman, T.C. Carter, C.L. Chambers, E.H. Gillam, K. Geluso, T.J. Weller, D.W. Burles, Q.E. Fletcher, K.J.O. Norquay, and J.R. Goheen. Environmental drivers of body size in North American bats. Functional Ecology 37:1020-1032. PDF

Brito, B.T., N.L. Bjornlie, M.D. Carling, and J.R. Goheen. Diet selection, commonness, and rarity in a pair of smooth-toothed pocket gophers. Journal of Mammalogy 104:915-928. PDF

Brown, B.R.P., J.R. Goheen, S.D. Newsome, R.M. Pringle, T.M. Palmer, L.M. Khasoha, and T.R. Kartzinel. Host phylogeny and functional traits differentiate gut microbiomes in a diverse natural community of small mammals. Molecular Ecology 32:2320-2334. PDF

Lujan, E, R. Nielsen, Z. Short, W.N. Watetu, L.M. Khasoha, T.M. Palmer, J.R. Goheen, and J.M. Alston. Symbiotic acacia ants drive nesting behavior by birds in an African savanna. Biotropica 55:1101-1105. PDF

Molina, F.J., J.A. Smith, E. Donadio, A.D. Middleton, J.N. Pauli, and J.R. Goheen. Food limitation reduces risk avoidance by prey, but does not increase kill rates in a simple predator-prey system. Ecosphere 14:e4701. PDF

Pringle, R.M., J.O. Abraham, T.M. Anderson, T.C. Coverdale, A.B. Davies, C.L. Dutton, A. Gaylard, J.R. Goheen, R.M. Holdo, M.C. Hutchinson, D.M. Kimuyu, R.A. Long, A.L. Subalusky, and M.P. Veldhuis. Impacts of large herbivores on terrestrial ecosystems. Current Biology 33:R584-R610. PDF

Tucker, M.A. and 172 coauthors. Behavioral responses of terrestrial mammals to COVID-19 lockdowns. Science 380:1059-1064. PDF Cover

Wells, H.B.M., R.D. Crego, J.M. Alston, S.K. Ndung'u, L.M. Khasoha, C.G. Reed, A.A. Hassan, S. Kurukura, J. Ekadeli, M. Namoni, P.S. Stewart, D.M. Kimuyu, A.A. Wolf, T.P. Young, T.R. Kartzinel, T.M. Palmer, J.R. Goheen, and R.M. Pringle. 2023. Wild herbivores enhance resistance to invasion by exotic cacti in an African savanna. Journal of Ecology 111:33-44. PDF Cover

2022

Alston, J.M., M.E. Dillon, D.A. Keinath, I.M. Abernethy, and J.R. Goheen. Daily torpor reduces the energetic consequences of habitat selection for a widespread bat. Ecology 103:e3677. PDF

Alston, J.M., C.G. Reed, L.M. Khasoha, B.R.P. Brown, G. Busienei, N. Carlson, T.C. Coverdale, M. Dudenhoeffer, M.A. Dyck, J. Ekeno, A.A. Hassan, R. Hohbein, R.P. Jakopak, B. Kimiti, S. Kurukura, P. Lokeny, A.M. Louthan, S. Musila, P.M. Musili, T. Tindall, S. Weiner, T.R. Kartzinel, T.M. Palmer, R.M. Pringle, and J.R. Goheen. Ecological consequences of large herbivore exclusion in an African savanna: 12 years of data from the UHURU experiment. Ecology 103:e3649. PDF Cover

Broekman, M.J.E., and 87 coauthors. Evaluating expert-based habitat suitability information of terrestrial mammals with GPS tracking data. Global Ecology and Biogeography 31:1526-1541. PDF

Hays, B.R., C. Riginos, T.M. Palmer, D.F. Doak, B.C. Gituku, N.J. Maiyo, S. Mutisya, S. Musila, and J.R. Goheen. Demographic consequences of mutualism disruption: browsing and big-headed ant invasion drive acacia population declines. Ecology 103:e3655. PDF

Levine, R.L., R.A. Smiley, B.R. Jesmer, B.A. Oates, J.R. Goheen, T.R. Stephenson, M.J. Kauffman, G.L. Fralick, and K.L. Monteith. Extending body condition scoring beyond measurable rump fat to estimate full range of nutritional condition for moose. Alces 58:91-99. PDF

2021

Coverdale, T.C., R.D. O’Connell, M.C. Hutchinson, A. Savagian, T.R. Kartzinel, T.M. Palmer, J.R. Goheen, D.J. Augustine, M. Sankaran, C.E. Tarnita, and R.M. Pringle. Large herbivores suppress liana infestation in an African savanna. Proceedings of the National Academy of Sciences 118:e2101676118. PDF Cover

Esmaeili, S. and 48 coauthors. Body size and digestive system shape resource selection by ungulates: a cross-taxa test of the Forage Maturation Hypothesis. Ecology Letters 24:2178-2191. PDF

Guy, T.J., M.C. Hutchinson, K.C.R. Baldock, E. Kayser, B. Baiser, P.P.A. Staniczenko, J.R. Goheen, R.M. Pringle, and T.M. Palmer. Large herbivores disassemble plant-pollinator networks. Current Biology 31:2964-2971. PDF

Jesmer, B.R., M.J. Kauffman, A.B. Courtemanch, S. Kilpatrick, T. Thomas, J. Yost, K.L. Monteith, and J.R. Goheen. Life-history theory provides a framework for detecting resource limitation: a test of the Nutritional Buffer Hypothesis. Ecological Applications 31:e02299. PDF

Kauffman, M.J., E. Aikens, S. Esmaeili, P. Kaczensky, A.D. Middleton, K.L. Monteith, T.A. Morrison, T. Mueller, H. Sawyer, and J.R. Goheen. Causes, consequences, and conservation of ungulate migration. Annual Review of Ecology, Evolution, and Systematics 52:453-478. PDF

Milligan, P., T. Martin, G. John, C. Riginos, J.R. Goheen, S. Carpenter, and T.M. Palmer. Invasive ants reduce carbon fixation for a foundational East African ant-plant. Ecology Letters 24:1052-1062. PDF

Oates, B.A., K.L. Monteith, J.R. Goheen, J.A. Merkle, G.L. Fralick, and M.J. Kauffman. Detecting resource limitation in a large herbivore population is enhanced with measures of nutritional condition. Frontiers in Ecology and Evolution 8:522174. PDF

Pietrek, A.G., J.R. Goheen, C. Riginos, N.J. Maiyo, and T.M. Palmer. Density dependence and the spread of invasive big-headed ants in an East African savanna. Oecologia 195:667-676. PDF

Wells, H.B.M., R.D. Crego, Ø.H. Opedal, L.M. Khasoha, J.M. Alston, C.G. Reed, S. Weiner, D.M. Kimuyu, T.P. Young, S. Kurukura, A.A. Hassan, T.R. Kartzinel, T.M. Palmer, R.M. Pringle, and J.R. Goheen. Experimental evidence that effects of megaherbivores on mesoherbivore space use are influenced by species’ traits. Journal of Animal Ecology 90:2510-2522. PDF Cover

2020

Hays, B.R., C. Riginos, T.M. Palmer, B.C. Gituku, and J.R. Goheen. Using photography to estimate above-ground biomass of small trees. Journal of Tropical Ecology 36:213-219. PDF

Jesmer, B.R., M.J. Kauffman, M.A. Murphy, and J.R. Goheen. A test of the Niche Variation Hypothesis in a ruminant herbivore. Journal of Animal Ecology 89:2825-2839. PDF

Noonan, M.J. and 78 coauthors. Effects of body size on estimation of mammalian area requirements. Conservation Biology 34:1017-1028. PDF

Palmer, T.M., C. Riginos, P.D. Milligan, B.R. Hays, A.G. Pietrek, N.J. Maiyo, S. Mutisya, B. Gituku, S. Musila, S. Carpenter, and J.R. Goheen. Frenemy at the gate: invasion by Pheidole megacephala facilitates a competitively subordinate plant ant in Kenya. Ecology 102:e03230. PDF

2019

Ali, A.H., R. Amin, J.S. Evans, M. Fischer, A.T. Ford, A. Kibara, and J.R. Goheen. Evaluating support for range-restoration practices by rural Somalis: a win-win for local livelihoods and conservation of the world’s most endangered antelope? Animal Conservation 234:82-89. PDF

Alston, J.M., B.M. Maitland, B.T. Brito, S. Esmaeili, A.T. Ford, B. Hays, B.R. Jesmer, F.J. Molina, and J.R. Goheen. Reciprocity in restoration ecology: when might large carnivore reintroduction restore ecosystems? Biological Conservation 234:82-89. PDF

Alston, J.M., I.M. Abernethy, D.A. Keinath, and J.R. Goheen. Roost selection by male northern long-eared bats in a managed fire-adapted forest. Forest Ecology and Management 446:251-256. PDF

Bowyer, R.T., M.S. Boyce, J.R. Goheen, and J.L. Rachlow. Conservation of mammals: threats, protected areas, and hunting. Journal of Mammalogy 100:923-941. PDF

Coverdale, T.C., I.J. McGeary, R.D. O'Connell, T.M. Palmer, J.R. Goheen, M. Sankaran, D.J. Augustine, A.T. Ford, and R.M. Pringle. Strong but opposing effects of associational resistance and susceptibility on defense phenotype in an African savanna plant. Oikos 128:1772-1782. PDF

Esmaeili, S., M.R. Hemami, and J.R. Goheen. Human dimensions of wildlife conservation in Iran: assessment of human-wildlife conflict in restoring a wide-ranging endangered species. PLOS One 14:e0220702. PDF

Gill, B.A., P.M. Musili, S. Kurukura, A.A. Hassan, W.J. Kress, M. Kuzmina, J.R. Goheen, R.M. Pringle, and T.R. Kartzinel. Plant DNA metabarcode library and community phylogeny for a semi-arid savanna in Kenya. Molecular Ecology Resources 19:838-846. PDF

Louthan, A., E. Valencia, D.J. Martins, T. Guy, J. Goheen, T. Palmer, and D. Doak. Large mammals generate both top-down effects and extended trophic cascades on floral-visitor assemblages. Journal of Tropical Ecology 35:185-198. PDF

Ng'weno, C.C., A.T. Ford, A.K. Kibungei, and J.R. Goheen. Interspecific prey neighborhoods shape risk of predation in a savanna ecosystem. Ecology 100:e02698. PDF

Ng'weno, C.C., S.W. Buskirk, N.J. Georgiadis, B.C. Gituku, A.K. Kibungei, L.M. Porensky, D.I. Rubenstein, and J.R. Goheen. Apparent competition, lion predation, and livestock production: can conservation value be enhanced? Frontiers in Ecology and Evolution 7:123. PDF

Noonan, M.J. and 55 coauthors. A comprehensive analysis of autocorrelation and bias in home range estimation. Ecological Monographs 89:e01344. PDF

Oates, B.A., J.A. Merkle, M.J. Kauffman, S. Dewey, M.D. Jimenez, J.M. Vartanian, and J.R. Goheen. Antipredator response diminishes during periods of resource deficit for a large herbivore: a test of the starvation-predation hypothesis. Ecology 100:e02618. PDF Cover

Otieno, T.O., J.R. Goheen, P.W. Webala, A. Mwangi, I.M. Osuga, and A.T. Ford. Human- and risk-mediated browsing pressure by sympatric antelope in an African savanna. Biological Conservation 232:59-65. PDF

2018

Ali, A.H., M.J. Kauffman, R. Amin, A. Kibara, J. King, D. Mallon, C. Musyoki, and J.R. Goheen. Demographic drivers of a refugee species: large-scale experiments guide strategies for reintroductions of hirola. Ecological Applications 28:275-283. PDF Cover

Coverdale, T.C., J.R. Goheen, T.M. Palmer, and R.M. Pringle. Good neighbors make good defenses: associational refuges reduce defense investment in African savanna plants. Ecology 99:1724-1736. PDF Cover

Goheen, J.R. A win-win for wildlife and ranching. Nature Sustainability 1:535-536. PDF

Goheen, J.R., D.J. Augustine, K.E. Veblen, D.M. Kimuyu, L.M. Porensky, T.M. Palmer, L.M. Porensky, R.M. Pringle, J. Ratnam, C. Riginos, M. Sankaran, G.K. Charles, A.T. Ford, A.A. Hassan, R. Jakopak, T.R. Kartzinel, S. Kurukura, A.M. Louthan, W.O. Odadi, T.O. Otieno, A.M. Wambua, H.S. Young, and T.P. Young. Conservation and management lessons from large-mammal manipulations in East African rangelands: KLEE, GLADE, and UHURU experiments. Annals of the New York Academy of Sciences 1429:31-49. PDF

Jesmer, B.R., J.A. Merkle, J.R. Goheen, E.O. Aikens, J.L. Beck, A.B. Courtemanch, M.A. Hurley, D.E. McWhirter, H.M. Miyasaki, K.L. Monteith, and M.J. Kauffman. Is ungulate migration culturally transmitted? Evidence of social learning from translocated animals. Science 361:1023-1025. PDF Cover

Louthan, A.M., R.M. Pringle, J.R. Goheen, T.M. Palmer, W.F. Morris, and D.F. Doak. Aridity weakens population-level effects of multiple species interactions in Hibiscus meyeri. Proceedings of the National Academy of Sciences 115:543-548. PDF

Koerner, S.E. and 72 coauthors. Change in dominance determines herbivore effects on plant biodiversity. Nature Ecology and Evolution 2:1925-1932. PDF

Tucker, M.A. and 114 coauthors. Global reductions in terrestrial mammalian movements in human-dominated landscapes. Science 359:466-469. PDF Cover

2017

Ali, A.H., A.T. Ford, J.S. Evans, D. Mallon, M.M. Hayes, J. King, R. Amin, and J.R. Goheen. Resource selection and landscape change reveal mechanisms suppressing population recovery for the world's most endangered antelope. Journal of Applied Ecology 54:1720-1729. PDF

Ford, A.T., S.J. Cooke, J.R. Goheen, and T.P. Young. Conserving megafauna or sacrificing biodiversity? Bioscience 67:193-196. PDF

Jesmer, B.R, J.R. Goheen, K.L. Monteith, and M.J. Kauffman. State-dependent behavior alters endocrine-energy relationship: implications for conservation and management. Ecological Applications 27:2303-2312. PDF

Long, R.A., A. Wambua, J.R. Goheen, T.M. Palmer, and R.M. Pringle. Climatic variation modulates the indirect effects of large herbivores on small-mammal habitat use. Journal of Animal Ecology 86:739-748. PDF

Ng'weno, C.C., N.J. Maiyo, A.H. Ali, A.K. Kibungei, and J.R. Goheen. Lions influence the decline and habitat shift of hartebeest in a semiarid savanna. Journal of Mammalogy 98:1078-1087. PDF

2016

Coverdale, T.C., T.R. Kartzinel, K.L. Grabowski, R.K. Shriver, A.A. Hassan, J.R. Goheen, T.M. Palmer, and R.M. Pringle. Elephants in the understory: opposing direct and indirect effects of consumption and ecosystem engineering by megaherbivores. Ecology 97:3219-3230. PDF

Ernest, S.K.M., G.M. Yennie, G. Allington, E.M. Christensen, K. Geluso, J.R. Goheen, M.R. Schutzenhofer, S.R. Supp, K.M. Thibault, J.H. Brown, and T.J. Valone. Long-term monitoring and experimental manipulation of a Chihuahuan desert ecosystem near Portal, Arizona (1977-2013). Ecology Data Paper 97:1082. PDF

Goheen, J.R. Book Review of Serengeti IV. Journal of Mammalogy 97:1001-1002. PDF

Pringle, R.M., K.M. Prior, T.M. Palmer, T.P. Young, and J.R. Goheen. Large herbivores promote habitat specialization and beta diversity of African savanna trees. Ecology 97:2640-2657. PDF

Tabak, M.A., S. Poncet, K. Passfield, J.R. Goheen, and C.M. Del Rio. The ghost of invasives past: rat eradication and the community composition and energy flow of island bird communities. Ecosphere 7:e01442. PDF

2015

Ford, A.T. and J.R. Goheen. Trophic cascades by large carnivores: a case for strong inference and mechanism. Trends in Ecology and Evolution 30:725-735. PDF

Ford, A.T. and J.R. Goheen. An experimental study on risk effects in a dwarf antelope. Journal of Mammalogy 96:918-926. PDF

Ford, A.T., J.R. Goheen, D.J. Augustine, M.F. Kinnaird, T.G. O'Brien, T.M. Palmer, R.M. Pringle, and R. Woodroffe. Recovery of African wild dogs suppresses prey density but does not trigger a trophic cascade. Ecology 96:2705-2714. PDF

Fraser, L.H. and 61 coauthors. 2015. Worldwide evidence of a unimodal relationship between productivity and plant species richness. Science 349:302-305. PDF

Tabak, M.A., S. Poncet, K. Passfield, J.R. Goheen, and C.M. Del Rio. Rat eradication and the resistance and resilience of passerine bird assemblages in the Falkland Islands. Journal of Animal Ecology 84:755-764. PDF

Young, H.S., D.J. McCauley, R. Dirzo, J.R. Goheen, B. Agwanda, C. Brook, E. Otarola-Castillo, A.W. Ferguson, S.N. Kinyua, M.M. McDonough, T.M. Palmer, R.M. Pringle, T.P. Young, and K.M. Helgen. Context-dependent effects of large-wildlife declines on small-mammal communities in central Kenya. Ecological Applications 25:348-360. PDF Cover

2014

Ford, A.T., J.R. Goheen, T.O. Otieno, L. Bidner, L.A. Isbell, T.M. Palmer, D. Ward, R. Woodroffe, and R.M. Pringle. Large carnivores make savanna tree communities less thorny. Science 346:346-349. PDF

Kartzinel, T.R., J.R. Goheen, G.K. Charles, E. DeFranco, J.E. Maclean, T.O. Otieno, T.M. Palmer, and R.M. Pringle. Plant and small-mammal responses to large-herbivore exclusion in an African savanna: five years of the UHURU experiment. Ecology Data Paper 95:787. PDF

Louthan, A.M., D.F. Doak, J.R. Goheen, T.M. Palmer, and R.M. Pringle. Mechanisms of plant-plant interactions: concealment from herbivores is more important than abiotic-stress mediation in an African savannah. Proceedings of the Royal Society Series B PDF

Pringle, R.M., J.R. Goheen, T.M. Palmer, G.K. Charles, E. DeFranco, R. Hohbein, A.T. Ford, and C. Tarnita. Low functional redundancy among mammalian browsers in regulating an encroaching shrub in an African savannah. Proceedings of the Royal Society Series B PDF

2013

Goheen, J.R., T.M. Palmer, G.K. Charles, K.M. Helgen, S.N. Kinyua, J.E. Maclean, B.L. Turner, H.S. Young, and R.M. Pringle. Piecewise disassembly of a large-herbivore community across a rainfall gradient: the UHURU experiment. PLOS One 8:e55192. Goheen and Pringle contributed equally to this piece. PDF

Louthan, A.M., D.F. Doak, J.R. Goheen, T.M. Palmer, and R.M. Pringle. Climatic stress mediates the impacts of herbivory on plant population structure and components of individual fitness. Journal of Ecology 101:1074-1083. PDF

Palmer, T.M., M.L. Stanton, T.P. Young, J.S. Lemboi, J.R. Goheen, and R.M. Pringle. A role for indirect facilitation in maintaining diversity in a guild of African acacia ants. Ecology 94:1531-1539. PDF

Young, H.S., D.J. McCauley, K.M. Helgen, J.R. Goheen, E. Castillo, T.M. Palmer, R.M. Pringle, T.P. Young, and R. Dirzo. Variation in plant community response to experimental and anthropogenic driven declines in large wildlife. Journal of Ecology 101:1030-1041. PDF

2012

Seifert, A.W., S.G. Kiama, M.G. Seifert, J.R. Goheen, T.M. Palmer, and M. Maden. Skin shedding and tissue regeneration in African spiny mice (Acomys). Nature 489:561-565. PDF

2011

Maclean, J.E., J.R. Goheen, D.F. Doak, T.M. Palmer, and T.P. Young. Cryptic herbivores mediate the strength and form of ungulate impacts on a long-lived savanna tree. Ecology 92:1626-1636. PDF

2010

Goheen, J.R. and T.M. Palmer. Defensive plant-ants stabilize megaherbivore-driven landscape change in an African savanna. Current Biology 20:1768-1772. Authors contributed equally to this piece. PDF Cover

Goheen, J.R., T.M. Palmer, F. Keesing, C. Riginos, and T.P. Young. Large herbivores facilitate savanna tree establishment through diverse and indirect pathways. Journal of Animal Ecology 79:372-382. PDF

Palmer, T.M., D.F. Doak, M.L. Stanton, J.L. Bronstein, E.T. Kiers, T.P. Young, J.R. Goheen, and R.M. Pringle. Synergy of multiple partners, including freeloaders, increases host fitness in a multispecies mutualism. 2010. Proceedings of the National Academy of Sciences 107:17234-17239. PDF

Thibault, K.M., S.K.M. Ernest, E.P. White, J.H. Brown, and J.R. Goheen. Long-term insights into consumer-resource dynamics in desert rodents: effects of community composition across guilds. Journal of Mammalogy 91:787-797. PDF

2009

Drever, M.C., J.R. Goheen, and K. Martin. Species-energy theory, pulsed resources, and the regulation of species richness: avian community dynamics during a mountain pine beetle outbreak. Ecology 90:1095-1105. PDF

2008

Ernest, S.K.M, J.H. Brown, K.M. Thibault, E.P. White, and J.R. Goheen. Zero-sum, the niche, and metacommunities: long-term dynamics of community assembly. American Naturalist 172:E257-E269. PDF

Palmer, T.M., M.L. Stanton, T.P. Young, J.R. Goheen, R.M. Pringle, and R. Karban. Putting ant-acacia mutualisms to the fire: Response. Science 319:1759-1761. PDF

Palmer, T.M., M.L. Stanton, T.P. Young, J.R. Goheen, R.M. Pringle, and R. Karban. Breakdown of an ant-plant mutualism follows the loss of large herbivores from an African savanna. Science 319:192-195. PDF Cover

2007

Goheen, J.R., T.P. Young, F. Keesing, and T.M. Palmer. Consequences of herbivory by native ungulates for the reproduction of a savanna tree. Journal of Ecology 95:129-138. PDF

Lusk, J.J., R.K. Swihart, and J.R. Goheen. Correlates of interspecific synchrony and interannual variation in seed production by deciduous trees. Forest Ecology and Management 242:656-670. PDF

Poe, S., J.R. Goheen, and E.P. Hulebak. Convergent exaptation and adaptation in solitary island lizards. Proceedings of the Royal Society Series B 274:2231-2237. PDF

Swihart, R.K., J.R. Goheen, S.A. Schnelker, and C.E Rizkalla. Testing the generality of patch and landscape-level predictors of tree squirrel occurrence at a regional scale. Journal of Mammalogy 88:564-572. PDF

2006

Goheen, J.R., E.P. White, S.K.M. Ernest, and J.H. Brown. Intra-guild compensation regulates species richness in desert rodents: Reply. Ecology 87: 2121-2125. PDF

2005

Goheen, J.R. and R.K. Swihart. Resource selection and predation of North American red squirrels in deciduous forest fragments. Journal of Mammalogy 86:22-28. PDF

Goheen, J.R., E.P. White, S.K.M. Ernest, and J.H. Brown. Intra-guild compensation regulates species richness in desert rodents. Ecology 86:567-573. PDF

2004

Goheen, J.R., F. Keesing, B.F. Allan, D. Ogada, and R.S. Ostfeld. Net effects of large mammals on Acacia seedling survival in an African savanna. Ecology 85:1555-1561. PDF

2003

DeWoody, Y.D., R.K. Swihart, B.A. Craig, and J.R. Goheen. Diversity and stability in communities structured by asymmetric resource allocation. American Naturalist 162:514-527. PDF

Goheen, J.R., G.A. Kaufman, and D.W. Kaufman. Effect of body size on reproductive characteristics of the northern grasshopper mouse in north-central Kansas. Southwestern Naturalist 48:427-431. PDF

Goheen, J.R. and R.K. Swihart. Food-hoarding behavior of gray squirrels and North American red squirrels in the central hardwoods region: implications for forest regeneration. Canadian Journal of Zoology 81:1636-1639. PDF

Goheen, J.R., R.K. Swihart, T.M. Gehring, and M.S. Miller. Forces structuring tree squirrel communities in fragmented landscapes: species differences in forest connectivity and carrying capacity. Oikos 102:95-103. PDF

Goheen, J.R., R.K. Swihart, and J.H. Robins. The anatomy of a range expansion: changes in cranial morphology and rates of energy extraction for North American red squirrels from different latitudes. Oikos 102:33-44. PDF

Swihart, R.K., T.C. Atwood, J.R. Goheen, D. Scheiman, K. Munroe, and T.M. Gehring. Patch occupancy of North American mammals: Is patchiness in the eye of the beholder? Journal of Biogeography 30:1259-1279. PDF

2002

Goheen, J.R., G.A. Kaufman, and D.W. Kaufman. Patterns of reproduction for western harvest mice in north-central Kansas. Prairie Naturalist 34:107-113.

2001

Post, D.M., T.S. Armbrust, E.A. Horne, and J.R. Goheen. Sexual segregation results in differences in content and quality of bison (Bos bison) diets. Journal of Mammalogy 82:407-413. PDF

Schmidt, K.A., J.R. Goheen, and R. Naumann. Incidental nest predation in songbirds: behavioral indicators detect ecological scales and processes. Ecology 82:2937-2947. PDF

Schmidt, K.A., J.R. Goheen, R. Naumann, R.S. Ostfeld, E.M. Schauber, and A.K. Berkowitz. Experimental removal of strong and weak predators: mice and chipmunks preying on songbird nests. Ecology 82:2927-2936. PDF